Fishes that live in areas where anoxic conditions can occur often cannot obtain enough oxygen via gills and have accessory respiratory structures that enable them to breath air. The electric eel, in addition to gills, has an extensive series of highly vascularized papillae in the pharyngeal region. The eels rise to the surface to gulp air, which diffuses across the papillae into the blood. Other fishes swallow air and extract oxygen through vascularized regions of the gut. (Note that developmentally, lungs begin as evaginations of the gut.) Anabantid fishes (ex. bettas you see in pet stores) have vascularized chambers in the rear of the head, called labyrinths, which act in a similar manner. Many fishes are facultative air breathers; that is, oxygen uptake switches from gills to accessory organs when oxygen in the surrounding water becomes low. Others, like the electric eel and the anabantids, are obligatory air breathers. The gills alone cannot meet the respiratory needs of the fish even if the surrounding water is saturated with oxygen. These fishes drown if they cannot reach the surface to breath air.